There are 3 Primordial Vaults in Borderlands 4. Defeating the Guardian bosses at the end of all 3 Primordial Vaults unlocks the Nothing Left to Guard trophy & achievement.
Each main region has one Primordial Vault (The Fadefields, Carcadia Burn, Terminus Range). Dominion region doesn’t have a Vault. You must find 3 Vault Fragment collectibles in each region to obtain the Vault Key. Then the Primordial Vault icon appears on the map. When you approach it, a pillar will come out of the ground where you can insert the key. Then a purple portal appears, go through it to enter the Primordial Vault. There will be a side quest at the start of each vault. Accept it, then a yellow ramp activates. Use the ramp to be catapulted into an arena where you must defeat waves of enemies. You must fight through 3 arenas, then you fight the Vault Guardian boss in the 4th arena. Defeating the boss gives you access to a permanent power-up.
Beating the Primordial Vaults at Ultimate Vault Hunter Rank 4 (unlocked after story) is needed to level up to Ultimate Vault Hunter Rank 5 (UVH5), which in turn unlocks the Gear Five trophy & achievement. After the story this can be tracked under Missions > Challenges > Ultimate Vault Hunter.
For the 9 Vault Fragments refer to Borderlands 4 All Vault Key Fragment Locations.
Video Guide
Primordial Vault 1: Arch of Inceptus
Located in The Fadefields > Dissected Plateau. On top of a hill. Only spawns after finding the 3 Vault Fragments in The Fadefields. When you approach, a stone pillar will come out of the ground where you can insert the Fadefields Vault Key. Then a purple portal appears, step through it to enter the Primordial Vault. On the inside you can accept a side quest, then fight through 3 arenas and beat the Vault Guardian boss at the end.
Primordial Vault 2: Arch of Radix
Located in Carcadia Burn > Grindstone of the Worthy. Found in the desert sand along a road. Only spawns after finding the 3 Vault Fragments in Carcadia Burn. When you approach, a stone pillar will come out of the ground where you can insert the Carcadia Burn Vault Key. Then a purple portal appears, step through it to enter the Primordial Vault. On the inside you can accept a side quest, then fight through 3 arenas and beat the Vault Guardian boss at the end.
Primordial Vault 3: Arch of Origo
Located in Terminus Range > The Low Leys. This is hidden inside a mountain tunnel. You must glide to it across a gap, see the 2nd map image below, the yellow character arrow indicates the starting point. The 3rd image shows where you must glide to. Then at the end of the tunnel you reach the vault. Only spawns after finding the 3 Vault Fragments in Terminus Range. When you approach, a stone pillar will come out of the ground where you can insert the Terminus Range Vault Key. Then a purple portal appears, step through it to enter the Primordial Vault. On the inside you can accept a side quest, then fight through 3 arenas and beat the Vault Guardian boss at the end.
For all other collectibles refer to Borderlands 4 Collectible Guide.
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